The specific epithet refers to Benzilan Town, the type locality of this species.

Thallus non-lobate, areoles dispersed to continuous, tightly adnate to the substrate; upper surface pruinose, off-white, partially with orangish tinge; apothecia cryptolecanorine to lecanorine; containing norstictic, cryptostictic, and connorstictic acids.

China • Yunnan Prov.: Deqin Co., Benzilan Town, 28°11'35.79"N, 99°21'07.65"E, Along Jinshajiang River side, alt. 2108 m, on rock, 19 August 2018, Li-Song Wang et al. 18-60350 (KUN-L 63843).

Thallus areolate, non-lobate, tightly adnate to the substrate. Areoles dispersed to continuous, central areoles irregular, 0.5–1 mm wide, marginal areoles usually larger than the center, 3–10 mm wide, cracked; distant simple areoles circular, 0.5–3 mm wide. Upper surface plane to slightly convex, pruinose, off-white, with orangish tinge. Upper cortex paraplectenchymatous, 30–60 µm thick, inspersed with brown to dark brown granules (soluble in K); epinecral layer gelatinous, 10–20 µm thick, with hyaline crystals and dark brown granules (insoluble in K). Algal layer 50–125 µm thick, discontinuous; photobiont chlorococcoid, cells 8–20 µm diameter. Medulla opaque, filled with gray-black granules (not or partially soluble in K). Lower cortex absent.

Apothecia common (1–4 per areole), orbicular, 0.3–1 mm diameter, initially cryptolecanorine, becoming lecanorine and elevated above areoles at maturity, not constricted at base; disc slightly concave to plane, matt, black, pruinose; apothecial margin indistinct when young, becoming prominent (0.05–0.15 mm wide) during development, receding with disc expansion. Exciple absent or narrow, not exceeding 50 μm. Epithecium, hymenium and subhymenial layers combined 175–200 µm high; epithecium 10–20 µm high, containing hyaline crystals; epihymenium with brown granules (soluble in K) and black granules (insoluble in K), N−; hymenium 100–120 µm high, hyaline, I+ blue; subhymenial layers 50–75 µm high, hyaline, I+ blue; algal layer sparse below hypothecium; paraphyses simple, septate, non-moniliform, with 1–3 uppermost cells wider than the basal cells, 4–6 μm wide (basal cells ca. 2 μm wide); asci clavate, Aspicilia-type, 8-spored, 70–80 × 18–25 µm; ascospores simple, hyaline, broadly ellipsoid, (10.0-)11.5–12.4–13.2(-14.0) × (8.0-)7.9–8.6–9.3(-10.0) µm (n = 36), l/w ratio (1.2-)1.3–1.4–1.5(-1.7), wall ca.1 µm thick. Pycnidia few, punctiform, plane to slightly convex, 0.05–0.2 mm diameter; ostiole brown to dark brown; conidia bacilliform, hyaline, 4–6 × 1 µm.

Cortex K+ yellow, P−; Medulla K+ yellow to orangish red, P+ yellow, C−, KC−; norstictic, cryptostictic and connorstictic acids detected in TLC.

Growing on the exposed calcareous rock. Currently only known from the town of Benzilan, Yunnan, China.

This species is similar to Lobothallia gangwondoana in having a non-lobate thallus with dispersed areoles. However, L. gangwondoana differs in its larger areoles, epruinose and slightly brownish-gray to cacao-gray upper surface (vs. pruinose, off-white with light orangish tinge in L. benzilanensis), epruinose apothecial disc (vs. pruinose), and in the lack of cryptostictic and connorstictic acids (Kondratyuk et al. 2020).

China • Yunnan Prov.: Diqing Tibetan Autonomous Prefecture, Shangri-La City, near Jinshajiang River in Benzilan Vil., 28°11'35.79"N, 99°21'07.65"E, alt. 2108 m, on rock, 19 August 2018, Li-Song Wang et al. 18-60354 (KUN-L 63847).

The epithet refers to the flat thallus of this species.

Thallus appressed to the substrate, areolate with a flat and radiately lobate margin; upper surface orangish gray, covered by white pruina; apothecia lecanorine, orbicular, adnate, discs brown with orangish pruina; mature pycnidia protruding.

China • Sichuan Prov.: Aba Tibetan and Qiang Autonomous Prefecture, Li Co., Putou Vil., 31°25'08.23"N, 103°04'10.85"E, alt. 2099 m, on rock, 4 September 2020, Li-Song Wang et al. 20-67554 (KUN-L 75731).

Thallus areolate with lobate margin, up to 5 cm across, closely appressed to the substrate, 0.3–1.5 mm thick. Areoles polygonal to suborbicular, (0.3–)0.5–1.2(–2.0) mm wide, interspaces between areoles 0.05–0.1 mm wide. Lobes flat, ca. 0.3 mm thick, elongate, 2–3(–3.5) mm long, 0.4–0.8(–1.0) mm wide at base, 0.6–1.5 mm wide at apex. Upper surface matt, orangish gray, covered by white pruina. Upper cortex paraplectenchymatous, even, 20–30 μm thick, with brown granules (soluble in K); epinecral layer 10–15 µm thick, with dense black granules (insoluble in K). Algal layer 75–125 µm thick, discontinuous; photobiont chlorococcoid, cells 8–20 µm diameter. Medulla opaque, filled with gray-black granules. Lower cortex absent.

Apothecia lecanorine, scattered, orbicular, (0.4–)0.8–2.0(–2.5) mm in diameter, adnate, constricted at base at maturity; disc plane to slightly convex, matt, brown with orangish pruina; apothecial margin entire, orange-gray, pruinose, persistent, 0.1–0.25 mm wide. Exciple narrow, widening to 20–60 μm in the uppermost part. Epithecium, hymenium and subhymenial layers combined 130–180 µm high; epithecium 10–15 µm high; epihymenium 10–20 μm high, with brown to dark brown granules (mostly soluble in K, leaving faint brown residue), N−; hymenium 75–100 µm high, hyaline, I+ blue; subhymenial layers 40–75 μm high, hyaline, I+ blue; algal layer discontinuous below hypothecium, 20–50 μm high; paraphyses simple, rarely anastomosed, septate, submoniliform, with 2–3 uppermost cells shorter and wider than the basal cells, 3–4 μm wide (basal cells ca. 2 μm wide); asci clavate, Aspicilia-type, hyaline, 8-spored, 60–70 × 20–25 µm; ascospores simple, hyaline, broadly ellipsoid to spherical, (9.0-)9.5–10.4–11.3(-12.0) × (7.0-)7.7–8.7–9.8(-10.0) µm (n = 49), l/w ratio (1.0-)1.0–1.2–1.4(-1.6), wall ca.1 µm thick. Pycnidia common, 0.2–0.5 mm diameter, distinctly protruding at maturity; ostiole brownish to dark brown; conidia bacilliform, hyaline, 4–6 × 1 µm.

Cortex K+ yellow to red, P−; Medulla K+ yellow to red, P+ yellow, C−, KC−; norstictic and connorstictic acids detected by TLC.

Saxicolous. Currently only known in Sichuan Prov., China.

Lobothallia lobulata resembles the new species in the flat and appressed thallus. However, L. lobulata is readily distinguished by its light gray thallus surface bearing lobules, epruinose apothecial disc and the absence of secondary metabolites (Zhang et al. 2024a).

China • Sichuan Prov.: Aba Tibetan and Qiang Autonomous Prefecture, Li Co., Putou Vil., 31°25'08.23"N, 103°04'10.85"E, alt. 2099 m, on rock, 4 September 2020, Li-Song Wang et al. 20-66561 (KUN-L 74736) • Rangtang Co., 32°17'39.37"N, 101°00'44.83"E, alt. 3535 m, on rock, 4 September 2020, Xin-Yu Wang et al. XY20-2551 (KUN-L 78906).

The epithet refers to the numerous pycnidia per areole of this species.

Thallus areolate, with radiate marginal lobes, pycnidia prominent, usually numerous pycnidia per areole, curved and elongate. This species colonizes diverse substrates, predominantly calcareous and siliceous rocks, with an occasional occurrence on a coniferous stump.

China • Qinghai Prov.: Qumalai Co., Qumahe Vil., 34°54'53.01"N, 94°46'17.29"E, alt. 4412 m, on rock, 17 September 2020, Xin-Yu Wang et al. XY20-3034 (KUN-L 79390).

Thallus adnate to the substratum, up to 5–7 cm across, centrally areolate, 2–4 mm thick, marginally lobate, 0.5–1 mm thick. Areoles (0.5–)0.7–1(–1.5) mm wide, angular to rounded, slightly constricted at base, interspaces between areoles (0.05–)0.1–0.2(–0.4) mm wide, surface flat to slightly convex, grayish-brown covered with white pruina. Lobes simple to irregularly branched, radiate, not overlapping, (1–)2–4(–5) mm long, (0.6–)1–1.5 mm wide, surface pale brown with thin pruina. Upper cortex paraplectenchymatous, 30–50 μm thick, with brown granules (soluble in K); epinecral layer 10–20 µm thick, gelatinous with dark granules (insoluble in K). Algal layer (75–)100–150(–250) μm thick, discontinuous; photobiont chlorococcoid, cells 8–16 μm diam. Medulla 0.5–1 mm thick, opaque, filled with gray-black granules (insoluble or partially soluble in K). Lower cortex absent.

Apothecia lecanorine, numerous, 1–3 per areole, (0.3–)0.8–2(–2.5) mm in diameter, clustered or scattered, orbicular to angular by pressure, not to slightly constricted at base; disc plane to slightly concave, black or brown, epruinose (rarely with thin pruina); apothecial margin entire, concolorous with thallus, slightly elevated, persistent but narrowing with disc expansion, 0.15–0.25 mm wide. Exciple narrow, 40–50 μm in the uppermost part. Epithecium, hymenium and subhymenial layers combined 125–150 µm high; epithecium 10–15 µm high; epihymenium 15–25 μm high, with brown (soluble in K) and pale brown (insoluble in K) granules, N−; hymenium 70–100 µm high, hyaline, I+ blue; subhymenial layers 40–80 μm high, hyaline, I+ blue, hypothecium 30–40 μm, algal layer discontinuous below hypothecium; paraphyses simple, septate, submoniliform to moniliform at the tips, with 2–5 uppermost cells shorter and wider than the basal cells, 4–6 μm wide (basal cells ca. 2 μm wide); asci clavate, Aspicilia-type, hyaline, 8-spored, 60–70 × 20–30 µm; ascospores simple, hyaline, broadly ellipsoid to ellipsoid, (10.0-)11.2–12.0–12.8(-13.0) × (7.0-)7.0–7.8–8.5(-9.0) µm (n = 38), l/w ratio (1.1-)1.4–1.6–1.8(-1.9), wall ca.1 µm thick. Pycnidia numerous, usually numerous pycnidia per areole, immersed, unilocular to multilocular, spot-like, 0.1–0.3 mm when young, shortly elongate to curved, 0.4–0.5 × 0.2–0.3 mm at maturity; ostioles dark brown to blackish; conidia bacilliform, hyaline, 5–6(–7) × 1 µm.

Cortex K+ yellow to orange-red, P–; medulla K+ yellow or orange-yellow, P– or P+ pale yellow (near the algal layer), C–, KC–; containing norstictic and connorstictic acids.

Saxicolous, occasionally growing on the coniferous stump. This species is distributed in Qinghai, Sichuan, and Xizang Prov., China.

Lobothallia hedinii is morphologically similar to L. polypycnidiata in general thallus appearance. However, the former differs in its gray to brownish red thallus, parallel lobes of consistent width, and its relatively sparse pycnidia. Chemically, although both species contain norstictic and connorstictic acids, they are distinguished by their medullary spot-test reactions: L. polypycnidiata exhibits P– or P+ pale yellow (only near the algal layer), whereas L. hedinii shows a distinct P+ yellow reaction. Furthermore, the species differ in reproductive development: L. polypycnidiata specimens consistently show scarce ascus maturation despite the apothecial density, while producing abundant pycnidia; in contrast, L. hedinii displays significantly higher ascus production but fewer pycnidia. Additionally, Lobothallia praeradiosa could be confused with L. polypycnidiata, but differs in its smoother upper surface, loosely adnate lobes with partially overlapping margins, orange-brown apothecial margin, and punctiform pycnidia (Kou et al. 2013; Paukov et al. 2019).

China • Qinghai Prov.: Yushu Tibetan Autonomous Prefecture, Qumalai Co., Qumalai Vil., 34°54'53.01"N, 94°46'17.29"E, alt. 4412 m, on rock, 17 September 2020, Xin-Yu Wang et al. XY20-753 (KUN-L 78517), Li-Song Wang et al. 20-68344 (KUN-L 76525), 20-67160 (KUN-L 75336) • Zhiduo Co., Zhiqu Vil., 34°00'44.49"N, 95°47'45.85"E, alt. 4171 m, on rock, 18 September 2020, Li-Song Wang et al. 20-68419 (KUN-L 76601), 20-68413 (KUN-L 76595), Xin-Yu Wang et al. XY20-808 (KUN-L 78572) • Zaduo Co., Angsai Vil., 32°50'52.90"N, 95°28'23.05"E, alt. 4051 m, on rock, 17 September 2020, Li-Song Wang et al. 20-67303 (KUN-L 75480), 20-67265 (KUN-L 75442) • Zhaqing Vil., 32°57'03.90"N, 95°05'29.65"E, alt. 4166 m, on rock, 18 September 2020, Li-Song Wang et al. 20-68582 (KUN-L 76764), 20-68583 (KUN-L 76765) • Ado Vil., 32°50'52.95"N, 95°28'24.98"E, alt. 4029 m, on dead coniferous wood, 20 September 2020, Li-Song Wang et al. 20-68672 (KUN-L 76855) • Haixi Mongol and Tibetan Autonomous Prefecture, Golmud Ci., 35°04'24.13"N, 94°07'46.04"E, alt. 4419 m, on rock, 17 September 2020, Li-Song Wang et al. 20-67134 (KUN-L 75310) • Sichuan Prov.: Garze Tibetan Autonomous Prefecture, Shiqu Co., Sexu Town, 33°07'53.16"N, 97°41'46.91"E, alt. 4214, on rock, 21 September 2020, Li-Song Wang et al. 20-68728 (KUN-L 76911) • Xizang Prov.: Langkazi Co., entrance to Karola Glacier, 28°53'54.24"N, 90°13'31.31"E, alt. 4758, on rock, 24 July 2019, Xin-Yu Wang et al. XY19-2271 (KUN-L 73043) • Chamdo Ci., Jiangda Co., Tongpu Vil., 31°38'30.46"N, 98°26'11.62"E, alt. 3948 m, on limestone, 23 September 2020, Li-Song Wang et al. 20-68878 (KUN-L 77062), 20-68840 (KUN-L 77024).

The epithet indicates that this species is similar to Lobothallia pruinosa in morphology.

This species is characterized by its tightly adnate and thin thallus, areolate center and shortly lobate margin; plane, white or brownish gray upper surface with fine granular pruina; cryptolecanorine apothecia; large ascospores (10.0-)11.8–13.3–14.8(-17.0) × (8.0-)8.5–9.3–10.0(-11.0) µm; and its long conidia (5–)6–8(–9) × 1 µm.

China • Xizang Autonomous Region: Shigatse Ci., Dingri Co., Zhaguo Vil., 28°35'23.26"N, 86°53'55.21"E, alt. 4320 m, on rock, 27 July 2019, Li-Song Wang et al. 19-64075 (KUN-L 68571).

Thallus tightly adnate to the substrate, up to 3 cm across, centrally areolate, 1–2 mm thick, marginally lobate, 0.3–0.8 mm thick. Areoles (0.3–)0.6–1(–1.5) mm wide, angular to rounded, flat, not constricted at base; interspaces between areoles 0.05–0.1 mm wide. Lobes radiate, short, plane, apices equal to slightly wider than base, 1–2 mm long, base 0.5–1 mm wide, apex 0.5–1.5 mm wide. Upper surface white or pale brownish gray, pruinose, brownish where pruina thin. Upper cortex paraplectenchymatous, 30–50 µm thick, containing dark brown granules (soluble in K); epinecral layer with dense grayish-black granules (insoluble in K), 10–40(–50) µm thick. Algal layer 70–100(–150) µm thick, discontinuous; photobiont chlorococcoid, (6–)8–20 µm diameter. Medulla 0.1–0.5 mm thick, opaque, filled with grayish-black granules (insoluble or partially soluble in K). Lower cortex absent.

Apothecia cryptolecanorine, common, grouped in the center, 1–3 per areole, angular, (0.1–)0.3–0.8(–1.0) mm in diameter; disc concave, brownish to black, matt, pruinose; apothecial margin indistinct, concolorous with the thallus. Exciple narrow, widening to 10–30(–50) μm in the uppermost part. Epithecium, hymenium and subhymenial layers combined (100–)125–150(–175) µm high; epithecium 5–15 µm thick, gelatinous; epihymenium 15–25 µm high, with dark brown to brownish granules (soluble in K), N–; hymenium (70–)90–100(–110) µm high, hyaline, I+ blue; subhymenial layers 50–75 µm high, hyaline, I+ blue; algal layer relatively continuous below hypothecium, 50–70 μm high; paraphyses submoniliform, simple, septate, with 1–3 uppermost cells shorter and wider than the basal cells, 4–6 μm wide (basal cells ca. 2 μm wide); asci clavate, Aspicilia-type, hyaline, 8-spored, 60–75 × 15–25 µm; ascospores simple, hyaline, broadly ellipsoid, rarely spherical, (10.0-)11.8–13.3–14.8(-17.0) × (8.0-)8.5–9.3–10.0(-11.0) µm (n = 86), l/w ratio (1.0-)1.2–1.4–1.6(-2.1), wall ca.1 µm thick. Pycnidia rare, punctiform, slightly convex, 0.05–0.1 mm diameter, ostiole brown to dark brown, conidia bacilliform, hyaline, (5–)6–8(–9) × 1 µm.

Cortex K+ pale yellow or orange, P–. Medulla K+ yellow to orange-red, P+ yellow, C–, KC–. Containing norstictic and connorstictic acids.

Saxicolous. Currently known only in Qinghai, Xizang Province, China.

Specimens of this new species were collected from Qinghai and Xizang Provinces, China, at elevations ranging from 3000 to 4500 m. The overall morphology of these specimens is uniform, with the exception of voucher 20-67831 (KUN-L 76010), which exhibits a thinner upper cortex composed of loosely arranged hyphae.

Phylogenetically, Lobothallia pruinosa and L. densipruinosa are the closest relatives of the new species. L. pruinosa can be distinguished by its lecanorine apothecia at maturity and broader lobes. L. densipruinosa differs in its whitish to light greenish gray and entirely pruinose thallus, apothecial disc dark olivaceous when wet and blackish when dry, and the absence of connorstictic acid (Kou et al. 2013; Ashraf et al. 2022). Lobothallia pakistanica resembles L. pseudopruinosa in its white, lobate thallus but differs in possessing larger central areoles, broader marginal lobes, and lacking of secondary metabolites (Zulfiqar et al. 2022).

Within Lobothallia, L. brachyloba, L. platycarpaL. pulvinata and this new species share an areolate thallus with lobate margins, a whitish to whitish-gray upper surface, and the presence of norstictic acid. Nevertheless, each can be distinguished by specific characters: L. brachyloba has sparse thalline and 1–7 apothecia per areole (Paukov et al. 2019); L. platycarpa possesses larger areoles, fewer and non-aggregated apothecia, and limited distribution in North Africa (Algeria) (Zulfiqar et al. 2022); And L. pulvinata exhibits a thinner thallus and upper cortex (10–15 µm), and lecanorine apothecia (Zulfiqar et al. 2023). Additionally, Lobothallia cheresina, L. controversa, and L. lacteola also have a tightly adnate, whitish to whitish–gray thallus, but these species differ from L. pseudopruinosa in their absent or indistinct marginal lobes and their different secondary metabolites.

Lobothallia pseudopruinosa. China • Xizang: Shigatse Ci., Dingri Co., Zhaguo Vil., G219, 28°35'09.99"N, 87°03'42.35"E, alt. 4306 m, on surface of weathered shale in open area, 16 June 2022, Li-Song Wang et al. 22-71222 (KUN-L 85846), Yan-Yun Zhang ZYY22-324 (KUN-L 81905), ZYY22-326 (KUN-L 81907) • Angren Co., 29°19'01.32"N, 87°01'59.60"E, alt. 4524 m, on rock, 19 July 2019, Li-Song Wang et al. 19-63621 (KUN-L 68115) • Qinghai Prov.: Yushu Tibetan Autonomous Prefecture, Qumalai Co., Qumahe Vil., 34°54'50.67"N, 94°46'20.74"E, alt. 4396 m, on limestone, 17 September 2020, Li-Song Wang et al. 20-68331 (KUN-L 76512) • Zeku Co., Maixiu Town, 35°15'53.57"N, 101°52'30.23"E, alt. 3143–3163 m, on rock, 8 July 2022, Xin-Yu Wang and Min Ai XY22-1089 (KUN-L 84907), XY22-1088 (KUN-L 84906), An-Cheng Yin and Han-Xiang Chen 22-72438 (KUN-L 87064) • Jiuzhi Co., NianBaoYuZe National Geopark, 33°13'57.57"N, 100°55'58.56"E, alt. 4063 m, on sandy soil, 8 September 2020, Li-Song Wang et al. 20-67831 (KUN-L 76010).

Lobothallia pruinosa. China • Nei Mongol: Bayanhot Town, Helan Mountain, alt. 1600 m, on rock, 17 August 2011, H.Y.Wang 20123575, 20123154, 20123282, 20123626 (SDNU) • Urat Rear Banner Co., alt. 1600 m, on rock, 19 August 2011, H.Y.Wang 20122917 (SDNU), D.B.Tong 20123276, 20123888 (SDNU) • Qinghai Prov.: Hualong Hui Ehtnic Autonomous Co., Yashiga Town, 36°03'52.78"N, 101°57'03.44"E, alt. 2002 m, on rock, 8 July 2022, Li-Song Wang et al. 22-73170 (KUN-L 87648).

The epithet refers to the beautiful morphology of the thallus of this species.

Thallus areolate with radiate lobes, upper surface white and pruinose; apothecia lecanorine, adnate and rounded, 1–2 apothecia per areole, apothecial margin distinct and persistent, disc plane to slightly convex, not to rarely with faint pruina, containing norstictic and stictic acids.

China • Xizang Autonomous Region: Shigatse Ci., Dingri Co., Zhaguo Vil., 28°35'24.49"N, 86°53'58.22"E, elev. 4293 m, on rock, 27 July 2019, Li-Song Wang et al. 19-66027 (KUN-L 70435).

Thallus tightly adnate to the substrate, 0.3–2 mm thick, centrally areolate, marginally lobate. Areoles 0.5–1.2 mm wide, angular to rounded, not constricted at base, interspaces between areoles 0.05–0.1 mm wide. Lobes radiate, short, 1–2.5 mm long, base width (0.3–)0.6–0.8(–1) mm, apex width 0.6–2 mm. Upper surface moderately convex, white, partially pale brown on marginal lobes, pruinose. Upper cortex paraplectenchymatous, even, 30–40 μm thick, containing brown granules (soluble in K); epinecral layer gelatinous, 10–20 μm thick. Algal layer 125–150 μm high, discontinuous; photobiont chlorococcoid, cells 6–18 µm diameter. Medulla 0.3–0.6 mm thick, opaque, filled with gray-black granules (insoluble in K). Lower cortex absent.

Apothecia lecanorine, common, 1–2 per areole, simple to grouped in the center, orbicular to slightly angular by pressure from adjacent apothecia, (0.3–)0.6–2 mm diameter, initially immersed, later sessile, not to slightly constricted at base; disc plane, matt, brownish-black to black, epruinose or faintly pruinose, shallowly fissured when overmature; apothecial margin entire, concolorous with thallus, well-developed and persistent, (0.05–)0.1–0.2(–0.25) mm. Exciple narrow, widening to 70–80 μm in the uppermost part. Epithecium, hymenium and subhymenial layers combined 125–160 µm high; epithecium 5–15 µm high, epihymenium 12.5–20 μm high, filled with dark brown to brown granules (mostly soluble in K), N–; hymenium 90–100 μm high, hyaline, I+ dark blue; subhymenial layers ca. 50 μm high, hyaline, I+ dark blue; discontinuous algal layer beneath hypothecium; paraphyses simple, septate, submoniliform at the tips, 1–4 uppermost cells shorter and broader than basal cells, 3–5 μm wide (basal cells 2–3 μm wide); asci clavate, Aspicilia-type, hyaline, 8-spored, 70–80 × 20–30 µm; ascospores simple, hyaline, ellipsoid to broadly ellipsoid, (10.0-)10.8–12.3–13.7(-16.0) × (8.0-)7.8–8.5–9.1(-10.0) µm (n = 30), l/w ratio (1.1-)1.3–1.5–1.7(-1.9), wall ca.1 µm thick. Pycnidia few, punctiform, flat to slightly convex, 0.15–0.2 mm diameter; ostiole brownish black; conidia bacilliform, hyaline, 6–8 × 1 µm.

Cortex K+ pale yellow or K–, P–; medulla K+ yellow to orange-red, P+ yellow or orange-red, C–, KC– or KC+ pale yellow; norstictic and stictic acids detected by TLC, some specimens additionally contain cryptostictic, connorstictic and constictic acids.

Saxicolous. Currently known from Xizang and Qinghai Provinces, China.

Lobothallia pseudopruinosa and L. pruinosa resemble L. pulchra in having an areolate thallus with radiate lobes and a white pruinose surface, but differ in their indistinct apothecial margin, distinctly pruinose discs, and the absence of stictic acid (Kou et al. 2013). Lobothallia wangii differs from L. pulchra in its predominantly cryptolecanorine apothecia, N+ epihymenium, and the absence of both marginal lobes and stictic acid. Lobothallia pakistanica and L. pulvinata also have a white thallus with central areoles and marginal lobes, but differ from L. pulchra in their thinner thalli, apothecia that are not constricted at base, and in the lack of stictic acid (Zulfiqar et al. 2022, 2023).

China • Xizang Autonomous Region: Tingri Co., Zhaguo Vi., 28°35'24.49"N, 86°53'58.22"E, alt. 4256–4299 m, on rock, 27 July 2019, Li-Song Wang et al. 19-65686 (KUN-L 70194), 19-65652 (KUN-L 70159) • Dagze Co., Bangdui Vi, 29°44'06.51"N, 91°24'55.29"E, alt. 3700 m, on rock, 16 July 2019, Li-Song Wang et al. 19-64597 (KUN-L 69093) • Qinghai: Yushu Tibetan Autonomous Prefecture, Zaduo Co., 32°52'43.64"N, 95°20'29.61"E, alt. 4074 m, on rock, 20 September 2020, Li-Song Wang et al. 20-67281 (KUN-L 75458) • Ado Vi., 32°52'43.35"N, 95°20'30.19"E, alt. 4080 m, on rock, 20 September 2020, Xin-Yu Wang et al. XY20-3169 (KUN-L 79525).

The specific epithet refers to the red reaction of medulla in C spot-test.

Thallus areolate with a lobate margin, upper surface white with pale brown tinge, pruinose; areoles raised when bearing apothecia or pycnidia; apothecia lecanorine, disc pruinose; pycnidia protruding; medulla C+ rose–red, contain gyrophoric acid.

China • Xinjiang Uygur Autonomous Region: Hejing Co., Baluntai Vil., 42°51'56.45"N, 86°27'09.16"E, alt. 2026 m, on rock, 02 July 2022, Xin-Yu Wang and Min Ai XY22-897 (holotype: KUN-L 84715).

Thallus closely adnate to substrate, up to 5 cm across, areolate, 1–4 mm thick, marginally lobate, 0.4–1 mm thick. Areoles (0.5–)0.8–2(–2.4) mm wide, contiguous, angular to rounded, non-constricted at base, interspaces between areoles 0.1–0.2 mm wide. Lobes slightly broader in the apex, 2–3 mm long, 0.7–1.4 mm wide at base, 0.6–2(–2.5) mm wide at apex. Upper surface convex, cracked, pruinose, white with pale brown tinge where pruina is thin. Upper cortex paraplectenchymatous, 20–40 μm thick, containing brown granules (soluble in K); epinecral layer composed of coarse plate-like crystals, gelatinous material, and gray-black granules (insoluble in K), 10–30 μm thick. Algal layer 75–150 μm high, discontinuous, interrupted by hyphae, 10–60 μm wide; photobiont chlorococcoid, cells 8–20 µm diameter. Medulla 0.5–1 mm high, filled with gray–black granules (generally insoluble in K). Lower cortex absent.

Apothecia lecanorine, common, usually 1 per areole, (0.3–)0.6–1.5(–2.0) mm diameter, rounded, sessile, slightly constricted at base when mature; disc slightly concave to plane, black, pruinose; apothecial margin entire, persistent, 0.1–0.25 mm wide, pruinose, concolorous with thallus. Exciple narrow, up to 40 μm wide. Epithecium, hymenium and subhymenial layers combined (150–)175–225(–250) µm high; epithecium 5–20 µm high; epihymenium 20–50 μm high, filled with dark brown granules (soluble in K), N–; hymenium 100–150 μm high, hyaline, I– or I+ weakly bluish in lower part; subhymenial layers 50–100 μm high, hypothecium 40–50 μm, hyaline, I+ blue; algal layer below hypothecium discontinuous, 50–60 μm high; paraphyses simple, septate, submoniliform at the tips, with 2–3 uppermost cells shorter and broader than basal cells, 3–6 μm wide (basal cells 2–3 μm wide); asci clavate, Aspicilia-type, hyaline, 8-spored, 70–85 × 20–30 µm; ascospores simple, hyaline, broadly ellipsoid, (10.0-)11.9–13.1–14.2(-15.0) × (8.0-)8.7–9.4–10.0(-11.0) µm (n = 48), l/w ratio (1.0-)1.3–1.4–1.5(-1.8), wall ca.1 µm thick. Pycnidia common, protruding, 0.1–0.25 mm in diameter, sometimes elongated, 0.5(–0.7) × 0.2 mm (length × width); ostiole brownish to dark brown; conidia bacilliform, hyaline, 5–6 × 1 µm.

Cortex: K+ pale yellow, P–; medulla: K+ pale yellow to orange or K–, P–, C+ rose–red, KC+ red to pale yellow; containing gyrophoric, norstictic, stictic, connorstictic and constictic acids.

Saxicolous. Currently known only from Xinjiang, China.

This species is readily recognized by its C+ rose–red medullary reaction, resulting from the production of gyrophoric acid. Lobothallia semisterilis resembles this species in its white to gray thallus and protruding pycnidia, but differs in being terricolous and lacking gyrophoric and stictic acids (Zhang et al. 2020). Lobothallia pruinosa shares the characteristic white pruina on both thallus and discs, but differs in its thinner thallus, punctiform pycnidia, and the presence of norstictic and connorstictic acids as the major secondary metabolite (Kou et al. 2013). Lobothallia pulchra is similar to L. rubra in possessing white thallus and lecanorine apothecia, but differs in having epruinose discs and lacking gyrophoric acid.

China • Xinjiang Uygur Autonomous Region: Hejing Co., Baluntai Town, 42°51'56.45"N, 86°27'09.16"E, alt. 2026–2046 m, on rock, 2 July 2022, An-Cheng Yin and Han-Xiang Chen 22-72301 (KUN-L 86927) • National Highway 216, 42°51'56.39"N, 86°27'09.31"E, alt. 2100 m, on limestone, 2 July 2022, Li-Song Wang et al. 22-72886 (KUN-L 87373).

The epithet refers to the areoles with a constricted and stipitate base.

Thallus brown, thick, centrally areolate, areoles with a stipitate base, marginally lobate, margin of lobes free from the substrate with granular pruina on the upper surface; apothecia lecanorine, constricted at the base, disc with conspicuous white pruina, apothecial margin persistent, epruinose, brown to orange-brown; hymenium pale brown.

China • Xinjiang Uygur Autonomous Region: Hejing Co., along road G218, 42°54'22.90"N, 86°17'36.95"E, alt. 2228 m, on rock, 01 July 2022, Li-Song Wang et al. 22-72871 (KUN-L 87358).

Thallus relatively loosely attached to the substrate, up to 4–6 cm across, centrally areolate, 3–6(–8) mm thick, marginally lobate, (0.5–)1–2 mm thick, free from the substrate. Areoles angular (0.6–1.2 mm wide) to elongated (1.5–2 × 0.6–1 mm wide), base constricted into a stipe, up to 4 mm long, interspaces between areoles 0.1–0.3 mm wide. Lobes long, simple to 2–3 branched, apices slightly wider than base, 3–5 mm long, 0.6–1.5 mm wide at base, 0.6–2.5 mm wide at apex. Upper surface flat to slightly convex, matt, brown, thinly pruinose, granular pruina present on the margin of lobes. Upper cortex paraplectenchymatous, even, 40–50 μm thick, inspersed with brownish and gray granules (soluble in K); epinecral layer gelatinous, 10–15 μm thick. Algal layer 75–150 μm high, discontinuous; photobiont chlorococcoid, cells 10–25 µm diameter. Medulla 0.6–3 mm high, filled with gray-black granules (generally insoluble in K). Lower cortex absent.

Apothecia lecanorine, common, 1–2 per areole, simple to grouped, orbicular to angular by pressure, (0.3–)0.5–1.5(–1.8) mm wide, constricted at base in maturity; disc concave to plane, brown to black, conspicuously pruinose; apothecial margin entire, persistent, epruinose, brown to orange-brown, 0.1–0.25(–0.3) mm wide. Exciple narrow, widening to 60–80 μm in the uppermost part. Epithecium, hymenium and subhymenial layers combined 125–175 µm high; epithecium 5–15 µm high, with hyaline plate-like crystals; epihymenium 12.5–37.5 μm high, filled with dark brown granules (soluble in K), N–; hymenium 100–120 μm high, pale brown (soluble in K), I+ dark blue; subhymenial layers 50–75 μm high, hypothecium 40–50 μm, pale brown (soluble in K), I+ dark blue; algal layer below hypothecium sparse or absent; paraphyses simple, septate, submoniliform at the tips, with 2–4 uppermost cells shorter and broader than basal cells, 3–5 μm wide (basal cells ca. 2 μm wide); asci clavate, Aspicilia-type, hyaline, 8-spored, 70–85 × 18–25 µm; ascospores simple, hyaline, broadly ellipsoid to ellipsoid, (9.0-)9.8–11.4–12.9(-15.0) × (7.0-)6.8–7.6–8.4(-11.0) µm (n = 30), l/w ratio (1.0-)1.3–1.5–1.7(-2.0), wall ca.1 µm thick. Pycnidia punctiform, plane to slightly convex, 0.1–0.25 mm diameter; ostiole brown to dark brown; conidia bacilliform, hyaline, 4–6 × 1 µm.

Cortex K+ pale yellow, P–; medulla K+ yellow, P+ orange, C–, KC–; containing norstictic, stictic, cryptostictic and connorstictic acids.

Saxicolous. Currently only known from Xinjiang Province, China.

Lobothallia praeradiosa resembles L. stipitata in having an areolate thallus with radiating lobes, a constricted apothecial base, orange-brown thallus and apothecial margins, and containing norstictic acid. However, the former differs in its epruinose disc, hyaline hymenium, and lack of stictic acid (Kou et al. 2013; Paukov et al. 2019; Ryan 2004). Lobothallia semisterilis shares narrowly elongated lobes with granular pruina on the margins, but differs in its terricolous habitat, white to gray thallus, apothecia-like pycnidia, and absence of stictic acid (Zhang et al. 2020).

China • Xinjiang Uygur Autonomous Region: Hejing Co., National Highway 218, 42°54'22.90"N, 86°17'36.95"E, alt. 2228 m, on limestone, 1 July 2022, Yan-Yun Zhang ZYY22-616 (AHUB 00470), ZYY22-617 (KUN-L 82191) • Baluntai Town, 42°54'36.56"N, 86°17'02.47"E, alt. 2171 m, on rock, 1 July 2022, Xin-Yu Wang and Min Ai XY22-887 (KUN-L 84705).

The epithet refers to the Chinese lichenologist Li-Song Wang, who collected the type specimen.

Thallus areolate without marginal lobes, upper surface white, flat; apothecia common, cryptolecanorine to lecanorine, disc black, pruinose, slightly concave to plane, often shallowly fissured at maturity; epihymenium N+ weakly greenish to green, containing norstictic and connorstictic acids.

China • Xizang Autonomous Region: Shigatse Ci., Dingri Co., along road G219, 28°35'10.18"N, 87°03'43.62"E, alt. 4311 m, on rock, 06 June 2022, Li-Song Wang et al. 22-71215 (KUN-L 85839).

Thallus tightly adnate to the substrate, 1–4 mm thick, areolate without marginal lobes, spreading in the field up to 25 cm across, the collected specimen is fragmented, up to 4 cm across. Central areoles commonly rounded, (0.2–)0.5–1(–1.6) mm diameter, not constricted at base, interspaces between areoles 0.1–0.2 mm wide; marginal areoles rounded to slightly elongated, 0.8–1.2 × 0.5–1.2 mm. Upper surface flat, white and pruinose. Upper cortex paraplectenchymatous, 30–50 μm thick, inspersed with pale gray granules (soluble in K), uppermost part filled with dark brown granules (partly soluble in K), 20–30 μm thick; epinecral layer gray dark, 20–50 μm thick. Algal layer 75–100 μm thick, mostly continuous, occasionally interrupted by fungal tissue (10–20 μm wide); photobiont chlorococcoid, cells 8–20 µm in diameter. Medulla 0.5–2 mm high, filled with gray-black granules (generally insoluble in K). Lower cortex absent.

Apothecia predominantly immersed and cryptolecanorine, rarely emergent and lecanorine, numerous, 1 to 3 per areole, (0.1–)0.3–1(–1.5) mm wide, simple or grouped, orbicular to angular by pressure; disc black, pruinose, slightly concave to plane, often shallowly fissured at maturity; apothecial margin entire, pruinose, concolorous with thallus, absent when young, appearing and gradually receding as the apothecium develops, (0.05–)0.1–0.2(–0.25) mm wide. Exciple narrow, widening to 40–80 μm in the uppermost part. Epithecium, hymenium and subhymenial layers combined 125–175 µm high; epithecium 5–15 µm high, with hyaline plate-like crystals; epihymenium 15–50 μm high, filled with brown to dark brown granules (dark soluble in K), N+ weakly greenish to green; hymenium (75–)100–125 μm high, hyaline, I+ pale blue to blue; subhymenial layers (40–)50–70(–100) μm high, hypothecium 20–60 μm, hyaline, I+ blue; algal layer below the hypothecium sparse or absent; paraphyses simple, occasionally anastomosing, septate, submoniliform, with 1–3(–4) uppermost cells shorter and broader, 4–6 μm wide (basal cells ca. 2 μm wide); asci clavate, Aspicilia-type, hyaline, 8-spored, 60–80 × 20–30 µm; ascospores simple, hyaline, broadly ellipsoid to ellipsoid, (10.0-)11.9–13.4–14.9(-17.0) × (7.0-)7.9–8.6–9.2(-10.0) µm (n = 89), l/w ratio (1.1-)1.4–1.6–1.8(-2.1), wall ca.1 µm thick. Pycnidia few, punctiform, plane to slightly convex, 0.1–0.2 mm diameter; ostiole dark brown; conidia bacilliform, hyaline, 5–7(–9) × 1 µm.

Cortex K+ pale yellow, P–; medulla K+ yellow to orange-red, P+ yellow, C–, KC–. Norstictic acid and connorstictic acids detected in TLC.

Saxicolous. Currently only known in Xizang, China.

Lobothallia controversa and L. cheresina resemble L. wangii but differ in having completely immersed apothecia and different secondary metabolites. Lobothallia controversa has two chemotypes (Roux et al. 2016): chemotype controversa exhibits no reaction to standard reagents (K, C, KC, P, I) but produces terpenoids, while chemotype reagens shows K+ and P+ reactions due to the presence of stictic acid complex along with terpenoids. Lobothallia cheresina has three chemotypes (Müller 1880; Roux 2012): chemotype cheresina lacks secondary metabolites; chemotype justii contains stictic acid as the main secondary metabolite; and chemotype microspora produces norstictic acid as the major compound. Lobothallia iqbalii and L. lacteola also possess a white, areolate thallus, and cryptolecanorine to lecanorine apothecia. However, Lobothallia iqbalii is distinguished by its plane to convex discs, raised and persistent apothecial margin, N– epihymenium, and absence of secondary metabolites or presence of only norstictic acid (Zulfiqar et al. 2022, 2023). Lobothallia lacteola differs from L. wangii in its thinner thallus, plicate margins, and absence of connorstictic acid (Paukov et al. 2019).

China • Xizang Autonomous Region: Shigatse Ci., Tingri Co., National Highway 219, 28°35'10.18"N, 87°03'43.62"E, alt. 4311 m, on schist, 16 June 2022, Yan-Yun Zhang ZYY22-324 (KUN-L 81905), ZYY22-331 (KUN-L 81912), ZYY22-334 (KUN-L 81915) • Zhaguo Vi.: 28°35'25.56"N, 86°53'56.03"E, alt. 4292 m, on rock, 27 July 2019, Li-Song Wang et al. 19-65657 (KUN-L 70164), 19-64063 (KUN-L 68559).

Thallus areolate with lobate margin, tightly adnate to the substrate, up to 3 cm across, 0.6–2 mm thick. Areoles (0.5–)0.8–1.5(–2) mm wide, irregular to suborbicular, not constricted at base, interspaces between areoles mostly 0.2–0.4 mm wide. Lobes short, simple to 2–3 branched, 2–3 mm long, 0.6–1.5 mm wide at the base, 1–3 mm wide at the apex, mostly 0.6–0.8 mm thick. Upper surface flat to slightly convex, light gray with brownish tinge, thinly pruinose. Upper cortex paraplectenchymatous, even, 30–40 μm thick, with brown granules (soluble in K); epinecral layer 5–20 µm thick, with brown to black granules (insoluble in K). Algal layer 100–125(–200) µm thick, discontinuous; photobiont chlorococcoid, cells 8–18 µm diameter. Medulla 0.4–1.6(–2) mm high, filled with gray-black granules. Lower cortex absent.

Apothecia cryptolecanorine (rarely elevated as lecanorine, not constricted at base), numerous, 1–8 per areole, orbicular to slightly angular, (0.1–)0.2–0.7(–1) mm diameter; disc slightly concave to flat, black, matt, epruinose to faintly pruinose; apothecial margin usually indistinct (when lecanorine, slightly elevated, 0.05–0.2 mm wide). Exciple narrow, widening to 20–30 μm in the uppermost part. Epithecium, hymenium and subhymenial layers combined 175–225 µm high; epithecium 5–15 µm high; epihymenium 15–40 μm high, with brown dark granules (mostly soluble in K), N−; hymenium 120–140 µm high, hyaline, I+ weakly bluish; subhymenial layers 60–75 μm high, hyaline, I+ blue; algal layer beneath hypothecium discontinuous; paraphyses simple, septate, submoniliform at the tips, with 2–3 uppermost cells shorter and wider than the basal cells, 3–5(–6) μm wide (basal cells ca. 2 μm wide); asci clavate, Aspicilia-type, hyaline, 8-spored, 60–70 × 15–22 µm; ascospores simple, hyaline, broadly ellipsoid, (9.0-)9.9–10.8–11.7(-12.0) × (7.0-)7.6–8.2–8.9(-10.0) µm (n = 41), l/w ratio (1.1-)1.2–1.3–1.4(-1.6), wall ca.1 µm thick. Pycnidia few, punctiform, plane to slightly convex, 0.05–0.2 mm diameter, occasionally elongate, 0.2–0.4 × 0.05–0.1 mm; ostioles dark brown; conidia bacilliform, hyaline, 5–6 × 1–1.5 µm.

Cortex K+ yellow to orange-red, P+ pale yellow; medulla K+ yellow to red, P+ yellow; containing norstictic acid.

Saxicolous. Currently known from the Altai mountains (Republic of Altai, Russia) and the Xinjiang Uygur Autonomous Region (China).

Lobothallia brachyloba is characterized by the areolate thallus with short marginal lobes, the light gray upper surface, the predominantly cryptolecanorine apothecia (1–8 per areole), and the presence of norstictic acid. Lobothallia crassimarginata resembles L. brachyloba in its gray areolate thallus, short marginal lobes and cryptolecanorine apothecia, but differs in typically bearing only one apothecium per areole, possessing a thick apothecial margin, and containing stictic acid (Kou et al. 2013; Paukov et al. 2019). Lobothallia benzilanensis shares the gray thallus and presence of norstictic acid with L. brachyloba, but can be distinguished by its dispersed to continuous thallus lacking marginal lobes and its fewer apothecia per areole.

China • Xinjiang Uygur Autonomous Region: Xinyuan Co., Nalati Town, 43°16'14.26"N, 84°30'08.89"E, alt. 1885 m, on limestone, 1 July 2022, Li-Song Wang et al. 22-72815 (KUN-L 87302).

Lecanora hedinii H. Magn. Lichens from Central Asia 1: 98 (1940).

China • Gansu Prov.: Eh-ma-ta-ch’üan, 1931, Bohlin 55a (S).

Thallus tightly adnate to the substrate, 3–6 cm across (up to 10 cm in intact field specimens), centrally areolate, 3–4 mm thick, marginally lobate, 0.4–1 mm thick. Areoles angular to irregular, not constricted at base, (0.4–)0.6–1(–1.8) mm wide, interspaces between areoles 0.05–0.2 mm wide. Lobes simple, rarely dichotomous, parallel-arranged, (1.5–)2–3(–4) mm long, 0.5–1.5(–2) mm wide. Upper surface plane to slightly convex, gray or pale brown to brownish red, pruinose, pruina usually thicker on the margin of areoles and lobes. Upper cortex paraplectenchymatous, 25–40 μm thick, containing brown granules (soluble in K); epinecral layer 10–15 µm thick, with dark granules (partly soluble in K). Algal layer 125–200 µm thick, discontinuous; photobiont chlorococcoid, cells 10–20 µm diameter. Medulla 0.5–2.5(–3) mm high, filled with gray-black granules (insoluble or partly soluble in K). Lower cortex absent.

Apothecia numerous, initially cryptolecanorine, becoming lecanorine at maturity, 1–3 per areole, adnate, orbicular to slightly angular by pressure, (0.2–)0.4–1.5(–2.2) mm wide; disc matt, brown to black, pruinose, concave to plane; apothecial margin entire, concolorous with thallus, 0.05–0.2 mm wide. Exciple narrow, widening to 50–80 μm in the uppermost part. Epithecium, hymenium and subhymenial layers combined 125–150 µm high; epithecium 10–15 µm high, with plate-like crystals; epihymenium filled with brown granules (soluble in K), N–; hymenium 75–100 µm high, hyaline, I+ blue; subhymenial layers 40–50 μm high, hyaline, I+ blue; algal layer beneath hypothecium discontinuous; paraphyses simple, septate, submoniliform at the tips, with 2–4 uppermost cells shorter and wider than the basal cells, 4–5 μm wide (basal cells ca. 2 μm wide); asci clavate, Aspicilia-type, hyaline, 8-spored, 65–75 × 20–30 µm; ascospores simple, hyaline, broadly ellipsoid to ellipsoid, (11.0-)12.5–13.5–14.5(-16.0) × (8.0-)8.4–9.2–9.9(-11.0) µm (n = 33), l/w ratio (1.2-)1.3–1.5–1.7(-1.9), wall ca.1 µm thick. Pycnidia common, punctiform when young, slightly protruding at maturity, 0.1–0.2(–0.5) mm in diameter; ostioles brown to black; conidia bacilliform, hyaline, 5–6 × 1 µm.

Cortex K+ yellow, P–; medulla K+ yellow to orange-red, P+ yellow, C–, KC–; containing norstictic and connorstictic acids, cryptostictic acid also detected in one specimen.

Saxicolous. Currently known from Gansu, Qinghai and Xizang Provinces, China.

The species was originally described from Gansu Province, China. Our integrated phenotypic and genotypic analyses confirmed its phylogenetic placement for the first time and extended its known distribution to Qinghai and Xizang. Although variable in thallus color, this species can be easily recognized by its areolate thallus with straight and parallel-arranged lobes, and production of norstictic and constictic acids. Its phylogenetic sister species, Lobothallia polypycnidiata, resembles L. hedinii but differs in having multiple pycnidia per areole, lobes with a slightly broader margin, and the P– or P+ pale yellow (only near the algal layer) medullary spot-reaction, contrasting with the distinct P+ yellow reaction in L. hedinii. Lobothallia praeradiosa shares production of norstictic and connorstictic acids with L. hedinii but differs in its curved and overlapping marginal lobes, gray or orange-brown upper surface, lecanorine apothecia, epruinose discs, and the orange-brown apothecial margins (Magnusson 1940; Kou et al. 2013; Paukov et al. 2019).

China • Gansu Prov.: Yumen City, Yu’erhong Vi., 39°39'29.84"N, 97°10'45.88"E, alt. 3044 m, on rock, 27 May 2018, Li-Song Wang et al. 18-59518 (KUN-L 62904), 18-59517 (KUN-L 62903) • Sunan Yugur Autonomous Co., Dahe Vi., 38°51'29.84"N, 99°38'55.16"E, alt. 2265 m, on sandstone, 6 July 2022, An-Cheng Yin and Han-Xiang Chen 22-72380 (KUN-L 87006) • Qinghai Prov.: Haixi Prefecture, Dulan Co., Xiangjia Vi., 36°00'53.98"N, 97°44'35.61"E, alt. 3036 m, on rock, 15 September 2020, Xin-Yu Wang et al. XY20-2969 (KUN-L 79324) • Dulan Co., S.B. Zhang ZSB24-1727 (KUN-L), Dulan-64 (KUN-L) • Golmud City, 35°53'06.05"N, 94°28'22.55"E, alt. 3665 m, on rock, 16 September 2020, Li-Song Wang et al. 20-68298 (KUN-L 76479), 20-68279 (KUN-L 76460) • Delingha City, S.B. Zhang Dacaidan-9; Hainan Prefecture, Gonghe Co., 36°46'21.59"N, 99°38'47.62"E, 3456 m, Li-Song Wang et al. 18-58220 (KUN-L 61713) • Tianjun Co., 37°38'24"N, 98°37'29"E, alt. 3581 m, on rock, 1 January 2023, S.B.Zhang and H.L.Kang ZSB23-410 (KUN-L 83735) • Yushu Tibetan Autonomous Prefecture, Zaduo Co., 32°52'43.73"N, 95°20'28.82"E, alt. 4083 m, on sandstone, 20 September 2020, Li-Song Wang et al. 20-68620 (KUN-L 76802) • Golog Tibetan Autonomous Prefecture, Gande Co., 33°50'29.51"N, 99°40'33.09"E, alt. 3986 m, on rocks, 13 September 2020, Li-Song Wang et al. 20-68152 (KUN-L 76332), 20-68147 (KUN-L 76327), S.B.Zhang ZSB24-1730 (KUN-L) • Xizang Autonomous Region: Sakya Co., Jiding Town, 29°12'01.18"N, 88°23'10.59"E, alt. 3941 m, on calcareous rocks, 14 June 2022, Li-Song Wang et al. 22-71169 (KUN-L 85793), 22-71155 (KUN-L 85779) • Qushui Co., Qushui Town, 29°23'03.77"N, 90°50'03.23"E, alt. 3580 m, on rocks, 17 July 2019, Li-Song Wang et al. 19-64640 (KUN-L 69137).