Maximum Likelihood (RAxML) tree of concatenated nrDNA ITS, nrDNA LSU and tef1-α sequences of representative species of the genus Tulostoma and the four newly-introduced species in the present study. Sequences obtained in this study are shown in bold blue. After the voucher number, the species and the country of origin are shown. Then, the type specimens are indicated. ML bootstrap support values (≥ 70) are shown before slashes and Bayesian posterior probabilities (≥ 0.90) are shown after slashes. Highlighted sections indicate affiliations to the four novel Tulostoma species: T. dunense, T. hungaricum, T. sacchariolens and T. shaihuludii. The illustrations exhibit basidiomata and basidiospore characteristics of the novel species. Tulostoma pulchellum (MJ7833) and T. striatum (Fritz) served as multiple outgroups. The scale bar indicates expected changes per site per branch.